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Darwin and Modern Science [59]

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a century, has become the starting-point for the new Swedish principle of selecting agricultural plants. It was the principle of single-ear sowing, instead of mixing the grains of all the selected ears together. By sowing each ear on a separate plot he intended not only to multiply them, but also to compare their value. This comparison ultimately led him to the choice of some few valuable sorts, one of which, the "Bellevue de Talavera," still holds its place among the prominent sorts of wheat cultivated in France. This variety seems to be really a uniform type, a quality very useful under favourable conditions of cultivation, but which seems to have destroyed its capacity for further improvement by selection.

The principle of single-ear sowing, with a view to obtain pure and uniform strains without further selection, has, until a few years ago, been almost entirely lost sight of. Only a very few agriculturists have applied it: among these are Patrick Shirreff ("Die Verbesserung der Getreide-Arten", translated by R. Hesse, Halle, 1880.) in Scotland and Willet M. Hays ("Wheat, varieties, breeding, cultivation", Univ. Minnesota, Agricultural Experimental Station, Bull. no. 62, 1899.) in Minnesota. Patrick Shirreff observed the fact, that in large fields of cereals, single plants may from time to time be found with larger ears, which justify the expectation of a far greater yield. In the course of about twenty-five years he isolated in this way two varieties of wheat and two of oats. He simply multiplied them as fast as possible, without any selection, and put them on the market.

Hays was struck by the fact that the yield of wheat in Minnesota was far beneath that in the neighbouring States. The local varieties were Fife and Blue Stem. They gave him, on inspection, some better specimens, "phenomenal yielders" as he called them. These were simply isolated and propagated, and, after comparison with the parent-variety and with some other selected strains of less value, were judged to be of sufficient importance to be tested by cultivation all over the State of Minnesota. They have since almost supplanted the original types, at least in most parts of the State, with the result that the total yield of wheat in Minnesota is said to have been increased by about a million dollars yearly.

Definite progress in the method of single-ear sowing has, however, been made only recently. It had been foreshadowed by Patrick Shirreff, who after the production of the four varieties already mentioned, tried to carry out his work on a larger scale, by including numerous minor deviations from the main type. He found by doing so that the chances of obtaining a better form were sufficiently increased to justify the trial. But it was Nilsson who discovered the almost inexhaustible polymorphy of cereals and other agricultural crops and made it the starting-point for a new and entirely trustworthy method of the highest utility. By this means he has produced during the last fifteen years a number of new and valuable races, which have already supplanted the old types on numerous farms in Sweden and which are now being introduced on a large scale into Germany and other European countries.

It is now twenty years since the station at Svalof was founded. During the first period of its work, embracing about five years, selection was practised on the principle which was then generally used in Germany. In order to improve a race a sample of the best ears was carefully selected from the best fields of the variety. These ears were considered as representatives of the type under cultivation, and it was assumed that by sowing their grains on a small plot a family could be obtained, which could afterwards be improved by a continuous selection. Differences between the collected ears were either not observed or disregarded. At Svalof this method of selection was practised on a far larger scale than on any German farm, and the result was, broadly speaking, the same. This may be stated in the following words: improvement in a few cases,
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