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Darwin and Modern Science [62]

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how the variations to be selected arise. They may arise slowly, from simple fluctuations, or suddenly, by mutations; in both cases natural selection will take hold of them, will multiply them if they are beneficial, and in the course of time accumulate them, so as to produce that great diversity of organic life, which we so highly admire.

Darwin has left the decision of this difficult and obviously subordinate point to his followers. But in his Pangenesis hypothesis he has given us the clue for a close study and ultimate elucidation of the subject under discussion.


V. HEREDITY AND VARIATION IN MODERN LIGHTS.

By W. BATESON, M.A., F.R.S.

Professor of Biology in the University of Cambridge.

Darwin's work has the property of greatness in that it may be admired from more aspects than one. For some the perception of the principle of Natural Selection stands out as his most wonderful achievement to which all the rest is subordinate. Others, among whom I would range myself, look up to him rather as the first who plainly distinguished, collected, and comprehensively studied that new class of evidence from which hereafter a true understanding of the process of Evolution may be developed. We each prefer our own standpoint of admiration; but I think that it will be in their wider aspect that his labours will most command the veneration of posterity.

A treatise written to advance knowledge may be read in two moods. The reader may keep his mind passive, willing merely to receive the impress of the writer's thought; or he may read with his attention strained and alert, asking at every instant how the new knowledge can be used in a further advance, watching continually for fresh footholds by which to climb higher still. Of Shelley it has been said that he was a poet for poets: so Darwin was a naturalist for naturalists. It is when his writings are used in the critical and more exacting spirit with which we test the outfit for our own enterprise that we learn their full value and strength. Whether we glance back and compare his performance with the efforts of his predecessors, or look forward along the course which modern research is disclosing, we shall honour most in him not the rounded merit of finite accomplishment, but the creative power by which he inaugurated a line of discovery endless in variety and extension. Let us attempt thus to see his work in true perspective between the past from which it grew, and the present which is its consequence. Darwin attacked the problem of Evolution by reference to facts of three classes: Variation; Heredity; Natural Selection. His work was not as the laity suppose, a sudden and unheralded revelation, but the first fruit of a long and hitherto barren controversy. The occurrence of variation from type, and the hereditary transmission of such variation had of course been long familiar to practical men, and inferences as to the possible bearing of those phenomena on the nature of specific difference had been from time to time drawn by naturalists. Maupertuis, for example, wrote "Ce qui nous reste a examiner, c'est comment d'un seul individu, il a pu naitre tant d'especes si differentes." And again "La Nature contient le fonds de toutes ces varietes: mais le hasard ou l'art les mettent en oeuvre. C'est ainsi que ceux dont l'industrie s'applique a satisfaire le gout des curieux, sont, pour ainsi dire, creatures d'especes nouvelles." ("Venus Physique, contenant deux Dissertations, l'une sur l'origine des Hommes et des Animaux: Et l'autre sur l'origine des Noirs" La Haye, 1746, pages 124 and 129. For an introduction to the writings of Maupertuis I am indebted to an article by Professor Lovejoy in "Popular Sci. Monthly", 1902.)

Such passages, of which many (though few so emphatic) can be found in eighteenth century writers, indicate a true perception of the mode of Evolution. The speculations hinted at by Buffon (For the fullest account of the views of these pioneers of Evolution, see the works of Samuel Butler, especially "Evolution, Old and New" (2nd edition)
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