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The Life and Letters-2 [209]

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had appeared before the 'Origin,' the author would have been canonised rather than anathematised by the natural theologians.) This is illustrated by a review in the "Literary Churchman", in which only one fault found, namely, that Mr. Darwin's expression of admiration at the contrivances in orchids is too indirect a way of saying, "O Lord, how manifold are Thy works!"

A somewhat similar criticism occurs in the 'Edinburgh Review' (October 1862). The writer points out that Mr. Darwin constantly uses phrases, such as "beautiful contrivance," "the labellum is...IN ORDER TO attract," "the nectar is PURPOSELY lodged." The Reviewer concludes his discussion thus: "We know, too that these purposes and ideas are not our own, but the ideas and purposes of Another."

The 'Edinburgh' reviewer's treatment of this subject was criticised in the "Saturday Review", November 15th, 1862: With reference to this article my father wrote to Sir Joseph Hooker (December 29th, 1862):--

"Here is an odd chance; my nephew Henry Parker, an Oxford Classic, and Fellow of Oriel, came here this evening; and I asked him whether he knew who had written the little article in the "Saturday", smashing the [Edinburgh reviewer], which we liked; and after a little hesitation he owned he had. I never knew that he wrote in the "Saturday"; and was it not an odd chance?"

The 'Edinburgh' article was written by the Duke of Argyll, and has since been made use of in his 'Reign of Law,' 1867. Mr. Wallace replied ('Quarterly Journal of Science,' October 1867. Republished in 'Natural Selection,' 1871.) to the Duke's criticisms, making some specially good remarks on those which refer to orchids. He shows how, by a "beautiful self-acting adjustment," the nectary of the orchid Angraecum (from 10 to 14 inches in length), and the proboscis of a moth sufficiently long to reach the nectar, might be developed by natural selection. He goes on to point out that on any other theory we must suppose that the flower was created with an enormously long nectary, and that then by a special act, an insect was created fitted to visit the flower, which would otherwise remain sterile. With regard to this point my father wrote (October 12 or 13, 1867):--

"I forgot to remark how capitally you turn the tables on the Duke, when you make him create the Angraecum and Moth by special creation."

If we examine the literature relating to the fertilisation of flowers, we do not find that this new branch of study showed any great activity immediately after the publication of the Orchid-book. There are a few papers by Asa Gray, in 1862 and 1863, by Hildebrand in 1864, and by Moggridge in 1865, but the great mass of work by Axell, Delpino, Hildebrand, and the Mullers, did not begin to appear until about 1867. The period during which the new views were being assimilated, and before they became thoroughly fruitful, was, however, surprisingly short. The later activity in this department may be roughly gauged by the fact that the valuable 'Bibliography,' given by Prof. D'Arcy Thompson in his translation of Muller's 'Befruchtung' (1883), contains references to 814 papers.

Besides the book on Orchids, my father wrote two or three papers on the subject, which will be found mentioned in the Appendix. The earliest of these, on the three sexual forms of Catasetum, was published in 1862; it is an anticipation of part of the Orchid-book, and was merely published in the Linnean Society's Journal, in acknowledgment of the use made of a specimen in the Society's possession. The possibility of apparently distinct species being merely sexual forms of a single species, suggested a characteristic experiment, which is alluded to in the following letter to one of his earliest disciples in the study of the fertilisation of flowers:]


CHARLES DARWIN TO J. TRAHERNE MOGGRIDGE. (The late Mr. Moggridge, author of 'Harvesting Ants and Trap-door Spiders,' 'Flora of Mentone,' etc.) Down, October 13 [1865].

My dear Sir,

I am especially obliged to you for your beautiful plates and letter-press;
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