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More Letters of Charles Darwin Volume I [205]

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was Secretary of the Anthropologische Gesellschaft of Munich, in which Society took place the discussion referred to in "Variation of Animals and Plants," I., 459, as originating Darwin's doubts on the whole question. The fresh evidence adduced by Kollmann as to the normal occurrence of a rudimentary sixth digit in Batrachians is Borus' paper, "Die sechste Zehe der Anuren" in "Morpholog. Jahrbuch," Bd. I., page 435. On this subject see Letter 178.), saying he was sorry that I had given up atavism and extra digits, and telling me of new and good evidence of rudiments of a rudimentary sixth digit in Batrachians (which I had myself seen, but given up owing to Gegenbaur's views); but, with re-growth failing me, I could not uphold my old notion.


LETTER 275. TO G.J. ROMANES.

(275/1. Mr. Romanes' reply to this letter is printed in his "Life and Letters," page 93, where by an oversight it is dated 1880-81.)

H. Wedgwood, Esq., Hopedene, Dorking, May 29th [1876].

As you are interested in pangenesis, and will some day, I hope, convert an "airy nothing" into a substantial theory, I send by this post an essay by Hackel (275/2. "Die Perigenesis der Plastidule oder die Wellenzeugung der Lebenstheilchen," 79 pages. Berlin, 1876.) attacking Pan. and substituting a molecular hypothesis. If I understand his views rightly, he would say that with a bird which strengthened its wings by use, the formative protoplasm of the strengthened parts became changed, and its molecular vibrations consequently changed, and that these vibrations are transmitted throughout the whole frame of the bird, and affect the sexual elements in such a manner that the wings of the offspring are developed in a like strengthened manner. I imagine he would say, in cases like those of Lord Morton's mare (275/3. A nearly pure-bred Arabian chestnut mare bore a hybrid to a quagga, and subsequently produced two striped colts by a black Arabian horse: see "Animals and Plants," I., page 403. The case was originally described in the "Philosophical Transactions," 1821, page 20. For an account of recent work bearing on this question, see article on "Zebras, Horses, and Hybrids," in the "Quarterly Review," October 1899. See Letter 235.), that the vibrations from the protoplasm, or "plasson," of the seminal fluid of the zebra set plasson vibrating in the mare; and that these vibrations continued until the hair of the second colt was formed, and which consequently became barred like that of a zebra. How he explains reversion to a remote ancestor, I know not. Perhaps I have misunderstood him, though I have skimmed the whole with some care. He lays much stress on inheritance being a form of unconscious memory, but how far this is part of his molecular vibration, I do not understand. His views make nothing clearer to me; but this may be my fault. No one, I presume, would doubt about molecular movements of some kind. His essay is clever and striking. If you read it (but you must not on my account), I should much like to hear your judgment, and you can return it at any time. The blue lines are Hackel's to call my attention.

We have come here for rest for me, which I have much needed; and shall remain here for about ten days more, and then home to work, which is my sole pleasure in life. I hope your splendid Medusa work and your experiments on pangenesis are going on well. I heard from my son Frank yesterday that he was feverish with a cold, and could not dine with the physiologists, which I am very sorry for, as I should have heard what they think about the new Bill. I see that you are one of the secretaries to this young Society.


LETTER 276. TO H.N. MOSELEY. Down, November 22nd [1876].

It is very kind of you to send me the Japanese books, which are extremely curious and amusing. My son Frank is away, but I am sure he will be much obliged for the two papers which you have sent him.

Thanks, also, for your interesting note. It is a pity that Peripatus (276/1. Moseley "On the Structure and Development of Peripatus capensis" ("Phil. Trans.
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